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Volume 80, Issue 1


Morpho-anatomical characteristics of Indian pangolin (Manis crassicaudata) from Potohar Plateau, Pakistan

Nausheen Irshad
  • Department of Wildlife Management, PMAS-Arid Agriculture University, Rawalpindi 46300, Pakistan
  • Other articles by this author:
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/ Tariq Mahmood
  • Corresponding author
  • Department of Wildlife Management, PMAS-Arid Agriculture University, Rawalpindi 46300, Pakistan
  • Email
  • Other articles by this author:
  • De Gruyter OnlineGoogle Scholar
/ Muhammad Sajid Nadeem
  • Department of Wildlife Management, PMAS-Arid Agriculture University, Rawalpindi 46300, Pakistan
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Published Online: 2015-02-04 | DOI: https://doi.org/10.1515/mammalia-2014-0179


We collected 10 live and two dead specimens of Indian pangolin (Manis crassicaudata) from Potohar Plateau to record their morpho-anatomical characteristics. The average body weight and length were recorded in three age classes: juvenile, sub-adult and adult. The maximum body weight, body length and tail length was 20 kg, 147.3 cm, and 85.7 cm, respectively. The total number of scales was 286±6.4, 271±24 and 294±11.0 in the juvenile, sub-adult and adult stages, respectively. The size of the scales varied on different body regions, with smallest being on the head and the largest on the trunk region. The number of scales on the head differed significantly from that on the trunk (p<0.01) and tail (p<0.05) regions. The number and size of the scales on the fore limbs differed significantly (p<0.05) from those on the hind limbs. The relative tongue length constituted 37% of the total body length of the adult specimens. The digestive tract constituted 7% of the relative body weight and 19%–23% of the relative body length, whereas relative length of the reproductive tract was 12%.

Keywords: anatomy; Indian pangolin; morphometery; Pakistan


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About the article

Corresponding author: Tariq Mahmood, Assistant Professor, Department of Wildlife Management, PMAS-Arid Agriculture University, Rawalpindi 46300, Pakistan, e-mail:

Received: 2014-12-19

Accepted: 2014-12-19

Published Online: 2015-02-04

Published in Print: 2016-01-01

Citation Information: Mammalia, Volume 80, Issue 1, Pages 103–110, ISSN (Online) 1864-1547, ISSN (Print) 0025-1461, DOI: https://doi.org/10.1515/mammalia-2014-0179.

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